The markings, which have a central line made by a heavy, plated tail, represent one of the largest invertebrate trackways found in the fossil record.

Dr Martin Whyte reports his discovery in central Scotland in Nature magazine.

"I knew it was a trackway as soon as I saw it - my main work is on dinosaur footprints - but it wasn't immediately obvious to me what sort of trackway it was," the Sheffield University researcher said.

"I could rule out reptiles and amphibians because I could see whatever it was, it had six legs," he told the BBC News Website.

"I've worked through the possibilities and I think it can only be Hibbertopterus."

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